Recently, the species Staphylococcus argenteus (silver staph) and Staphylococcus schweitzeri have been described (1). Both are more closely related to S. aureus than is Staphylococcus simae, which was previously regarded as the closest relative of S. aureus. S. aureus, S. argenteus and S. schweitzeri may be regarded as the “S. aureus complex”. S. argenteus is a globally distributed human pathogen with similar virulence properties to S. aureus, and has also been identified in an African bat population. S schweitzeri to date has been identified in African non-human primates, once in a human carrier in Gabon, and the same African bat population shown to harbour S. argenteus. Here we report large scale comparative genomics of S. argenteus and S. schweitzeri. The genome sequences of 180 S. argenteus and 24 S. schweitzeri isolates were determined. The principal findings of bioinformatic analyses were: 1. As foreshadowed in analysis of a smaller collection of isolates, the species are highly divergent throughout the great majority of their core genomes, supporting their classification as separate species, 2. S. argenteus and S. schweitzeri are phylogenetic sister groups within the S. aureus complex, despite the close resemblance between S. aureus and S. argenteus as regards known host range and virulence properties, 3. The population structure of the sequenced S. argenteus isolates is discontinuous, consisting of seven highly conserved clonal complexes, indicating multiple genetic bottlenecks in the recent past, 4. There have been more instances of core genome horizontal transfer between S. aureus and S. schweitzeri than between S. argenteus and either of the other two species, and 5. S. argenteus is rapidly acquiring accessory genome elements characteristic of S. aureus. Our current model is that multiple cross species transfer events within the last 50 years have introduced S. argenteus into humans.